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Chapter
1: Introduction
evo01_introduction.wpd
evo01.pdf
Study guide
Chapter
2: Darwin and the evidence for evolution
evo02_evidence.wpd
evo02.pdf
Study guide
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Chapter
3: Darwinian natural selection
evo03_selection.wpd
evo03.pdf
Study guide
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Chapter
4: Mutation and genetic variation
evo04_genetics.wpd
evo04.pdf
Study guide
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Chapter
5: Mendelian genetics in populations I: selection and mutation as mechanisms
of evolution
evo05_hardyweinberg.wpd
evo05.pdf
Study guide
Answers to sample problems
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Chapter
6: Mendelian genetics in populations II: migration, drift, non-random mating
evo06_migration_etc.wpd
evo06.pdf
Study guide
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Several sites offer models of genetic drift. Genetic
Drift Simulation is a pencil-and-paper simulation from the University
of Arizona, good for teachers. Genetic
Drift , from the University of Connecticut, lets you plug in allele
frequencies and population size and graphs the results. Jon Herron
has developed some excellent simulations of drift and other processes.
Download the AlleleA1
software and exercises and enjoy.
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The effects of small populations on extinction risk are of major interest
to conservation biologists. Introduction
- ALEX introduces you to some of the simulation software used to assess
extinction risk based on a variety of ecological parameters using an approach
called Population Viability Analysis.
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The Florida Panther is a classic example of the effects of small population
size on inbreeding and inbreeding depression. Read a good review
at the Florida
Panather Net site (check out the link to "Threats").
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The Conservation Breeding Specialist
Group , a division of the IUCN, is an international conservation organization
whose mission is "to assist conservation of threatened animal and plant
species through scientific management of small populations in wild habitats,
with linkage to captive populations where needed." This site offers
some interesting information on how that's done.
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Chapter
7: Quantitative genetics
evo07_quantgen.wpd
evo07.pdf
measuring selection illustration
Study guide
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Many folks have critiqued The Bell Curve since its publication in
1994. Some good resources are Bell
Curve , a review article by Arthur S. Goldberger and Charles F. Manski
from the Department of Economics
at The University of Wisconsin-Madison ; The
Bell Curve Flattened - Subsequent research has seriously undercut the claims
of the controversial best seller. by Nicholas Lemann from Slate magazine
and BELL CURVE: Anatomy
of an Analysis by Claudia Krenz , a statistician who has re-analyzed
the data from the book.
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Heritability is an important concept in animal breeding. Read Heritability
by John Pollack, Ph.D., Cornell University and G2910
Heritability and Its Use in Animal Breeding by John W. Massey and Dale
W. Vogt, Department of Animal Sciences, University of Missouri-Columbia
for more information.
-
Evolution
of Polymorphisms in seedcrackers describes ongoing research on these
birds, along with a bibliography and some great photos, by the folks at
the Center for Tropical Studies at The University of California, San Francisco.
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NNZ
- Goldenrod Gall Fly is a neat short article on gall flies. From
its contents, you should be able to develop some good hypotheses for the
causes of stabilizing selection in this critter. Teachers should
check out Lab
6 - Gall Size , from Jonathan Brown, Ph.D., at Grinnell College for
a good lab exercise.
-
If you're interested in the study of genetic linkage (especially as it
applies to the study of human genetics and genetic disease), check out
genetic
linkage analysis , the home page of the Laboratory of Satistical Genetics
at Rockefeller University
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Chapter
8: Studying adaptation: evolutionary analysis of form and function
evo08_adaptation.wpd
evo08.pdf
Study guide
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The strength of natural selection and the role of constraints, trade-offs,
and historical contingency have been the subject of debate among evolutionary
biologists since the late 1970's. Two major players in this debate
have been Richard Dawkins (arguing on the side of the pre-eminence of natural
selection) and Stephen J. Gould (arguing on the side of important limits
to selection in the forms of constraint and contingency). Read Stephen
J. Gould’s original argument from his classic paper “The Spandrels
of San Marco and the Panglossian Paradigm” at An
adaptationist programme , a site hosted by the America Academy
for the Advancement of Science. Then read some of their ongoing debates
at The "G-Files"
from John Catalano’s excellent website and follow up with Presidential
Lectures: Stephen Jay Gould: Commentary: Thurtle , a discussion of
their ongoing disagreement by Phillip Stevens Thurtle of Stanford University.
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DesertUSA 2000 is an on-line magazine
about US Deserts features good discussions of the adaptations of plants
and animals to survival
in arid environments. Bearing in mind that the statement that a trait
is an adaptation is really a hypothesis, how would you test the hypotheses
presented in this articles?
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Department of
Plant Population Biology: NIOO-Centre of Terrestrial Ecology: research
program and themes presents summaries of ongoing research on phenotypic
plasticity in plants. Some of the descriptions include results of
preliminary studies; see if you can make predictions about the outcomes
of the future studies as described on the site.
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CAIC
Homepage is the website for the software package Comparative Analysis
by Independent Contrasts, developed by Andy Purvis and Andrew Rambau to
calculate phylogenetically independent contrasts for comparative studies.
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Chapter
9: Sexual selection
evo09_sexual_selection.wpd
evo09.pdf
Study guide
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Sexual selection and the evolution of mating systems are major research
topics at a number of institutions. Read descriptions of current
research projects from The
Department of Biological and Environmental Sciences at the University
of Jrvaskyla; Dr.
Frederick Widemo at the Department of Animal Ecology, Evolutionary
Biology Centre in Upsala; and Dr.
Matt Persons at Susquehana University . Each of these pages provides
background and theory as well as information about the study organisms;
see if you can use this information to devise methods for testing the hypotheses
of interest to the researchers.
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Males’
seeming handicaps advertise a suitor’s worth , an article from US News
describes the “handicapped male” hypothesis, developed by Amotz Zahavi
in the mid 1970's. How does this hypothesis fit into the breakdown
of hypotheses for female choice presented in the text? Another hypothesis,
presented as an alternative to the “good genes” hypotheses is described
at A
New Look at Darwinian Sexual Selection ; is this hypothesis really
novel?
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Population
density, social pathology, and behavioral ecology
By Jim Moore from the Anthropology Department at The University of
California, San Diego, this article from the journal Primates presents
an alternative to the sexual selection hypothesis for male infanticide
in two species of mammals.
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Nat'l Academy Press, Biodiversity II: (1996), 18
Comparative Behavioral and Biochemical Studies of Bowerbirds and the Evolution
of Bower-Building . This is a review article by Dr. Gerald Borgia from
the Department of Biology, University of Maryland, on the evolution of
a novel type of male ornamentation – decorated bowers built by male bowerbirds
of Australia.
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Chapter
10: Kin Selection and Social Behavior
evo10_kinselection.doc
evo10.pdf
Study guide
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Chapter
12: Mechanisms of speciation
evo12_speciation.wpd
evo12.pdf
Study guide
Animations
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What
is a Species, and What is Not? In this paper from the journal The Philosophy
of Science, Mayr explains the biological species concept in the context
of the larger question of whether or not species exist in nature.
He also offers his critique of alternative species concepts, including
the typological (morphological), evolutionary, and phylogenetic species
concepts.
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At
this site, Jeff Groth of the Department of Ornithology at the American
Museum of Natural History discusses species concepts in the context of
a specific research question about the specific status of crossbills. Crossbill
Natural History is a good place to start your review.
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Learn more about red wolves at Red
wolf, Canis rufus, U.S. Fish & Wildlife Service and play with on-line
mapping of a population in Algonquin State Park at Red
Wolves / Database .
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Observed
Instances of Speciation From the Talk.Origins archive, this paper discusses
different species concepts and processes of speciation, and provides an
overview (with references) of instances in which speciation has been observed,
either in the field or in laboratory populations. Its “sister page”,
Some
More Observed Speciation Events provides additional references.
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The Union of Concerned Scientists is
an organization of scientists engaged in a variety of political issues
with a strong basis in science. One issue of major concern is the
potential consequences of new uses of biotechnology, including the use
of genetically engineered crop plants. For a good overview of the
potential problems, check out their Biotechnology
- UCS page.
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Chapter
13: Reconstructing evolutionary trees
evo13_phylogeny.wpd
evo13.pdf
Study guide
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Chapter
14: The origin of life and Precambrian evolution
evo14_abiogenesis.doc
evo14.pdf
Study guide
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Chapter
15: The Cambrian explosion and beyond
evo15_cambrian.doc
evo15.pdf
Study guide
Ediacaran
Fossils of Canada This site, from the Miller Museum of Geology, Queen's
University, Canada, describes the significance of the Ediacaran fauna and
provides pictures of representative fossils.
Burgess Shale
fossils Andrew MacRae, at the University of Calgary, provides descriptions
of the Burgess Shale site as well as pictures of a variety of its fossils
and links to additional information.
The Cambrian
Period This page gives an overview of the Cambrian Period and provides
links to more information on its stratigraphy, tectonics, early life, and
more. From Science Daily magazine, read Large
Gene Study Questions Cambrian Explosion , an article describing the
molecular evidence challenging the timing of the Cambrian Explosion.
Punctuated
Equilibrium at Twenty Dr. Donald Prothero reviews the hypothesis of
punctuated equilibrium in this article from the journal Skeptic.
BBC
Evolution Weekend: Extinction Files - Mass Extinctions
Part of the BBC's excellent evolution web site, this page links you
to summary information about each of the "Big Five" mass extinction events.
The
Great Mystery: Background from UC Berkeley's Museum of Paleontology
discusses the end-Cretaceous mass extinction.
IUCN - The World Conservation Union The
World Conservation Union website offers a huge variety of information about
biodiversity conservation, including a searchable on-line version of the
Red Book of sensitive, threatened and endangered species.
Food and Agriculture Organization of the
United Nations
This site provides information about the state of important global
resources. Look at FAO - Forestry
- Home Page for data on the world's forests.
Center for Biodiversity
and ConservationThe American Museum of Natural History offers this
on-line version of Humans
and Other Catastrophes: Perspectives on Extinction, a report on the
Center for Biodiversity and Conservation's 1997 symposium at AMNH. Written
by conservationist and author Curt Meine, the report gives an overview
of extinction episodes, including the current "Sixth Extinction," the almost
certain human causation of recent extinctions, and some possible conservation
strategies for the future. For more information on anthropogenic extinctions,
see the list of news releases and other resources provided by David Ulansey
at Mass Extinction
Underway
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Chapter
16: Human evolution
evo16_humans.doc
evo16.pdf
evo_16briefhumans.wpd (Fall
2001)
evo16b.pdf (Fall 2001)
Study guide
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The web offers a wealth of sites with information related to human evolution.
One of the most comprehensive lists of sites (including links to images
of hominid fossils) are Jim
Foley’s Fossil Hominids: the evidence for human evolution .
-
Becoming Human: Paleoanthropology,
Evolution and Human Origins is an awesome site that includes an on-line
documentary of human evolution and interactive exhibits that allow users
to explore everything from basic evolutionary theory to profiles of the
members of the hominid family tree. It requires a fast connection,
but is well worth the visit.
-
Fossil Evidence for Human
Evolution in China is from Dennis Etler at UC Berkeley and focuses
specifically on human evolution in China, with links to a variety of resources
including fossil images and research publications. One of these is
The
African Emergence and Early Asian Dispersals of the Genus Homo,
an on-line version of an American Scientist article by Roy Larick and Russell
L. Ciochon that addresses, among other things, the African emergence and
multiregional hypotheses.
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Forensic Anthropology
is
part of the Osteointeractive web site, a fantastic resource for learning
about osteology, one of the underpinnings of paleoanthropology. The
site includes information on human osteology, paleopathology, and histology
as well as forensic anthropology.
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Handprint : Ancestral
Lines is a non-technical but good site that brings together information
about the hominid fossil record, tools and tool use, dispersal routes,
and more in an attractive, accessible format.
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Language
Acquisition This book chapter by Steven Pinker discusses some of his
ideas about the evolution of language.
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Kerry S. Kilburn,
Ph.D
Department
of Biological Sciences
Old
Dominion University
Norfolk, VA 23529 |
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