Plant Structure & Function: Roots

 Read the section in the text which deals with roots. That part of a plant that absorbs water and mineral from the soil and transports it to the
stem is called the root. Most roots of a plant are located under the soil and anchor the plant in this
location. However some plants such as orchids have aerial roots which absorb water from the air.
Roots are also used by plants to store food (starch) for future use by the plant. Roots may
classified according to morphology and site of origin.
 
 

Root Types Involving  Morphology

     Tap Roots : one large main root and several smaller secondary roots; examples –
      carrots, beets

     Fibrous Roots: many roots of the same size; example grass
 
 

 Root Types Involving Origin:

      true roots : roots which develop from seed
 

      adventitious roots : roots which develop from other plant parts: stem, leaves,
                                       buds

Prop roots are a type adventitious root

To learn the structures in a root you must be able to identify the varies tissues located in a cross section and a longitudinal section of a root. By examining the longitudinal section of a root you can see the relationship between the three areas of embryonic tissues (primary meristem) and the mature tissue located in a mature stem.

Examine Figures Handed Out In Lecture
 

 Longitudinal Section of Root.
 

Apical meristem - cell division occurs -  All cells located in the root are formed by
                               the  process  of mitosis which occurs in the apical meristem

Root cap - The root cap protects these delicate cells from being crushed as the root
                   tip grows through the soil.

Zone of cell elongation - The newly formed cells will undergo elongation above the
                                          zone of cell division  thus pushing the root apex through the
                                          soil.

Zone of maturation - The cells above the zone of elongation will mature and
                                       differentiate into specific tissues depending on their location
                                       in the root.
 
 
 

             Three  primary meristems  (protoderm, ground meristem and procambium)

Protoderm develops into the epidermis

Ground Meristem develops into the cortex

Procambium develops into vascular  tissue (xylem and phloem)

Examine Figure Handed Out In Lecture

      Epidermis has extensions called root hairs which absorb water and minerals

    Root hairs are unicellular  epidermal cells located only at the tips (beginning of
     the region of maturation) of all roots. They increase the  surface  area of the root
     tip for  water absorption. Most of the water absorbed by roots enters through the
    root hairs.  Root hairs are found in an area of approximately four inches  above
    the tip of the root.

  Cross Section Of Root:

Examine Figure Handed Out In Lecture

     Epidermis - prevents water loss

     Cortex - made up of parenchyma cells - function to store starch

      Endodermis (single ring of cells around the vascular tissue) -  provides selective
      absorption of mineral from the exterior to the xylem. It performs this task by
      forcing  water (minerals) to pass through the living cell membrane and not
      through cell wall by diffusion. Because of  the involvement of the cell membrane
      mineral must pass through by active transport. Active  transport allows for
      selective absorption. Thus, for minerals to move from the cortex to the vascular
      tissue they are screened by the membrane of the endodermis. The way the
      endodermis forces water to move through the living part is that there is a layer
      of wax (Casparian Strip) around the top, side, and bottom edges with the front
      and back open. Water can not move through this wax.

Examine Figure Handed Out In Lecture
 

Apoplast tissue of the root- water and minerals will move through root through nonliving parts of cell until the endodermis is encountered.

Symplast tissue of the root – water and minerals may move through living parts of the cell (slow process). Movement through endodermis must be symplastic. This allows for active transport and selectivity.

Pericycle -  retains the potential to become meristematic. In the older part of the root the pericycle   tissue forms secondary or lateral roots.  These roots  develop at a forty  five degree angle from the root it originates.

Phloem - Conducts photosynthates, located in the arms of the xylem tissue
 

Xylem - Conducts water, center of root

Protophloem/ Metaphloem

Protoxylem/ Metaxylem
 

Review  differences between monocots and dicots as to examples and cotyledons.
 

Transverse section of monocot roots

Strands of xylem and phloem are distributed in ground tissue.

Pith present

Mycorrhizae    Symbiotic relationship  of soil fungi and roots.

                  Do not move beyond the endodermis
 
 

Root Nodules for Nitrogen Fixation

Refer to figures in text.

        Rhizobium             bacteria that infects cortex with an infection thread and
                                        stimulates the cortex cells to undergo mitosis to form root
                                        nodule

                                        Bacteria cells fill plant cells forming  bacteriods

                                        Nitrogen Fixation    N2      converted into   NH4
 
 

Haustoria                    parasitic plants form these modified roots
 

Refer to figures in text.
 

Indicate from which primary meristem each of specific tissues listed above are formed (epidermis,
root hairs, cortex, vascular tissue - xylem and phloem, endodermis, pericycle and  vascular
cambium).
 
Primary Meristems  Tissues
protoderm
ground meristem
procambium

Click to view apical meristem http://www.mancol.edu/science/biology/plants_new/anatomy/rootder1.html
Click to view root dermal tissue  http://www.mancol.edu/science/biology/plants_new/anatomy/rootder2.html
Click to view ground tissue  http://www.mancol.edu/science/biology/plants_new/anatomy/rootgrnd.html
 

In your notes draw a cross section of a root at the region of differentiation or maturation and label
the following tissues: epidermis, cortex, endodermis, pericycle, primary xylem, primary phloem.

Using the text write a function for the tissues listed above.
 
 
Tissues Function
epidermis
cortex
endodermis
pericycle
primary xylem
primary phloem

Can you identify the various cell types which makeup the various tissues listed above?
 
 
 
Tissues Cell Types - parenchyma, sclerenchyma, collenchyma, epidermal, sieve tube, vessel element
epidermis
cortex
endodermis
pericycle
primary xylem
primary phloem